Let me just throw this out there. slugs were never at any point fertile. Fertilization happens when the snake ovulates or shortly after. So slug never saw genetic material from either parent. The more likely idea is that infertile eggs were "Super spiders" didn't make it past the first cellular division. However because we don't truly understand reptile DNA and the associated mutations of it we wont be able to say for sure that the morphs are determined at the time of the first division or even later on. it's more likely that the mutation kicks in later in the embryo's development since its things like pigment and what not that are effected. With all of that said I've long said that it's naive to think that the only thing these mutations effect physiologically is skin pigment, We see kinking, cleft jaws, small eye's wobble's, duck bills ect ect. All of those this are changes to the anatomy of the animal.
Still so much to learn and discover