Stress, which affects a number of systems in snakes, stuck shed, dehydration, increased risk for respiratory illnesses, and some research has suggested that even if you don't see any signs of these, long term low humidity can have a negative affect on liver and kidney function. Think about it: if your natural environmental needs say you should keep a certain level of hydration, but something is preventing you from getting that, you'll end up pretty miserable after a while, wouldn't you? Same with your snake. They live in borrows where humidity is a bit higher than out in the open. This is the environment they've adapted for, and the environment you should strive to recreate.
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