*With the possible exception of the Blackhead Spider.
While I'm not sure this one counts, because it changes the optics so much, if it is indeed free (or even largely free) of wobble . . . It might lend some evidence to the idea that the spider wobble is not in the brain per se, but an issue of the inner ear created by pigment distribution during development.
Like what Oxy said above, insufficient pigment in the cells of the inner ear affect balance and mess up the snake's little gyro stabilizers, so to speak. Sort of similar to the way inadequate pigments in the cilia create problems in dogs.
Sorry. Nerd moment over
(Disclaimer: I don't have Spiders. I like them, I've just never seen a spot in the collection for them.)