This was actually suggested by another breeder. He seems to think there's another gene going on and he was going to consult someone else, but I haven't heard back from him. Doesn't look like the Lesser has yellow belly in her but I don't know the markers and haven't bothered to look because I personally don't think other genes are going on in them. They'll be shedding this week/weekend and I'll see if her eyes change.
¯\_(ツ)_/¯