I've always used (and seen it used) to indicate a breeding. So a het albino X het toffee would produce a 1:3 phenotypic ratio of toffinos and normals (66% poss het for either albino or toffee). However it sometimes depends on context.
NERD actually stumbled on a very convenient genetic term to describe what we're seeing when two "compatible" mutant allelic traits combine to produce a unique phenotype: Compound heterozygous.http://www.newenglandreptile.com/ind...cts-like-super
NERD lists all of the following as compound heterozygous:http://www.newenglandreptile.com/ind...uper-balls/als
As you can see, the definition fits perfectly with the Toffino / Candino phenomenon... as well as several others that have created difficulties for ball python terminology.
For example:
Gg = gravel (dominant gene/ heterozygous)
Yy = yellowbelly (dominant gene/ heterozygous)
GY = highway (compound heterozygous)
Tt = het toffee (recessive gene/ heterozygous)
Aa = het albino (recessive gene/ heterozygous)
ta = Toffino (compound heterozygous)