I agree with the above, when she is removed from the cage she could be thinking it's food time. I feed mine in their enclosures but I'm not sure if you have a particular reason for feeding this way?
You say you handle her at least twice a day? I would cut this down to once a day for around 10-30 minutes. These are animals who tolerate being handled, they do not enjoy it as such. I would recommend lessening this to once a day unless digesting or shedding where it's advised not to handle.
A large 10 degree gradient per end of the enclosure isn't great for us to judge that you have the right husbandry; can you narrow this down for us?
Also anything else you can think that you may have changed in regards to her? Anything in the enclosure etc?
Also, you say "heat mat on warm side, hide on cold" you do have a hide at both ends don't you?
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