Snakes can suffocate, but the tubs are not air tight against the top of the rack slots, so I wouldn't worry too much about that.
I just find that without air holes my humidity get up around 88% with a small water bowl. With air holes it stays right at 60%, give or take a little for ambient room humidity.
Also, RetiredJedi and I did an informal experiment, and both found that our top tubs hold a little more humidity than the lower 3. Not sure why that happens, but it is of note.
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