Well you would need both chromosomes, simply as a matter of pairing and the animal being 2n. But the genes on the ball 15 need not necessarily be identical to the genes on the burm 15. As in this example. So, while ball tyr may occupy position XYZ on ball 15 That same position on burm 15 would hold an aminoacyltransferase...
In this example it would have to be that way. The hybrid animal would have no way to "acquire" a wild type gene at the corresponding locus on the chromosome of the non-species parent. Basically what you would have is a animal that is homozygous for T- but heterozygous at each locus.Or you think that an animal with one T- on the 15 and one T- on the 7 wouldn't present melamine?
BUt as I said, this example is just an off the cuff extreme. If I had to bet odds I would guess that both ball and burm carry their tyr on the same chromosome and at roughly the same position.