Rats have a very high rate of metabolism- they need food AND water at all times. Were they crowded in the box- maybe overheated too? Can't tell you why only one was eaten- but from now on, try to be more prepared when holding them for so long. Young rats are like teenage humans- hungry all the time. They were probably not immediate family to each other, so they may have been squabbling some too. Rats are normally quite social & not inclined to eat each other- that's a desperation move though- that was WAY too long without food (& water?).
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