I'm talking out of memory from reading around... Breeding too young, over breeding, and obesity can cause slugs. Over breeding, some females need to take a season off once every 2~3 seasons. Another common cause I've seen is keeping the temps too high for the female or breeding male, since too high temps damage viable sperm. Stress wouldn't necessarily produce slugs, but the female may reabsorb and not lay at all.
A small but mature female at a healthy body condition wouldn't necessarily produce slugs just for being small, just smaller clutches.
I'm also trying to find out whether producing slugs could be sometimes genetic?
I'm going to have my first clutch this year, and I've been scouring the internet.
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