Question Alex:
A saccule is intricately linked to balance and equilibrium - correct?
Therefore if someone has proved that the spider genetics result in a deformation to the inner ear, it should be easy to find - correct?
Or does it sound like someone's attempt to sound scholarly to explain an unknown genetic snafu? To be clear, this idea of an inner ear abnormality has been tossed around for a few years, but like many other things, it has gained credibility just by being repeated often. I know of no studies done.......and I would hazard to say none exist.
FWIW: I have the Hartline pdf somewhere on my hard drive. It says pretty much the same thing that your link does.