Excellent article!
As this sort of thing has been found in egg-laying lizards and birds, it seems likely to be able to occur in egg-laying snakes.
A single egg cell develops inside each follicle in the ovary. Ovulation occurs when the egg cell leaves the follicle. Fertilization occurs after ovulation.
Male spider x female normal produces both spider and normal offspring. So spider sperm can get to the eggs. But are there fewer spider offspring from this sort of mating than from a normal male x female spider mating? Anyone collected any statistics? For that matter has anyone collected statistics to see if there are the same, fewer or more spiders than normals from such matings? If there are statistically less than 50% spiders from spider x normal matings, it would be consistent with the lethal homozygous spider theory.