I'm not an expert by any means, but I believe they are referring to a gene that is expressing itself within the mutation but has not been singled out by itself or if it is it looks normal and needs to be paired up with the original gene in order to express itself again. IDK. Dont know much about hidden gene womas, never heard of a hidden gene spider, and the hidden gene that was acts with the lessers is the daddy gene, het daddys look normal and need to be paired with a lesser for the gene to be expressed and to make more platy daddys. Essentially it is assumed that their is another "hidden" gene expressing itself within a certain mutation causing a "different" look than what the gene would normally look like. Hopefully someone else can explain this better and correct my amateur lacking explaination.