Rich stated he's seen normals from Lesser X Lesser pairing that had bug eyes, which suggests that bug-eyes does not have to be paired with the Lesser gene to express, and that it's a separate genetic defect.
Can anyone else confirm? This is rather important...keeping track of bloodlines should allow bug-eyes to be bred out if it is s separate recessive gene.
It's possible Rich uncovered a rare recessive bug-eye gene that is actually not related to the typical lesser BEL bug eye problem, of course, if no one else has ever seen this happen. If so, then bug-eyed BELs probably don't have a genetic defect (in most cases), they're just expressing an issue with the morph itself.