I think NERD has the most prolific "mega combos", although I know some others are also working on such things. Which has the most? I have no idea...but maybe someone does.
The trouble when you get into these "mega combos" is that it gets harder and harder to identify with certainty exactly which traits have been passed along. Someone with an eye like Kevin's can pick out minute details of each one...but even he doesn't know what, exactly, some of his combos are.